| CPC C07K 14/54 (2013.01) [A61P 35/00 (2018.01); C07K 14/7155 (2013.01); C07K 16/2818 (2013.01); A61K 38/00 (2013.01); C07K 2319/30 (2013.01)] | 25 Claims |

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1. A prodrug comprising a human IL-21 agonist polypeptide, a masking moiety, and a carrier moiety, wherein the human IL-21 agonist polypeptide comprises SEQ ID NO: 1, or comprise an amino acid sequence identical to SEQ ID NO: 1 but for one or more mutations selected from D18A/K, Q19K, and E109R (numbering according to SEQ ID NO: 1), the masking moiety comprises consists of an extracellular domain (ECD) of a human IL-21 receptor α or γ subunit,
the human IL-21 agonist polypeptide is fused to the carrier moiety, and
the masking moiety is fused to the human IL-21 agonist polypeptide or to the carrier moiety through a cleavable peptide linker.
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