| CPC C12N 15/11 (2013.01) [C12N 9/22 (2013.01); C12N 15/907 (2013.01); C12N 2310/20 (2017.05)] | 3 Claims |
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1. A method of producing a genetically modified eukaryotic cell having a base substitution, comprising:
(a) preparing a composition for base editing comprising
(i) an adenosine base editor (ABE) or cytosine base editor (CBE),
wherein the ABE is selected from the group consisting of apolipoprotein B editing complex 1 (APOBEC1) and activation-induced deaminase (AID), the CBE is tRNA-specific adenosine deaminase (tadA);
(ii) Cas 9 nickase (nCas9); and
(iii) an extended guide RNA (ggX20) hybridizable with a target sequence,
wherein the ggX20 comprises a sequence of 20 nucleotides of the target sequence, and two additional non-complementary guanines (G) in front of 20 nucleotides in the 5′ direction from protospacer adjacent motif (PAM),
wherein the target sequence is selected from HEK2, HBB, and FANCF;
(b) introducing the composition into a eukaryotic cell,
wherein the extended guide RNA hybridizes with the target sequence,
wherein the nCas9 makes a single stranded nick,
wherein the ABE converts an adenosine to a guanine at position 18 of the target sequence in the 5′ direction from the PAM;
wherein the CBE converts a cytosine to a thymine at position 20, 21, or 23 of the target sequence in the 5′ direction from the PAM, and
wherein the base substitution does not cause a double stranded break.
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