| CPC A61K 39/255 (2013.01) [A61K 39/17 (2013.01); A61P 31/20 (2018.01); C12N 15/86 (2013.01); A61K 2039/5254 (2013.01); A61K 2039/5256 (2013.01); A61K 2039/552 (2013.01); A61K 2039/70 (2013.01)] | 12 Claims |

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1. A recombinant herpesvirus of turkeys (rHVT) that comprises a first heterologous nucleic acid and a second heterologous nucleic acid;
wherein the first heterologous nucleic acid is located in a first nonessential site in the rHVT genome, and the second heterologous nucleic acid is located in a second nonessential site in the rHVT genome;
wherein the first nonessential site is the UL54.5 site and the second nonessential site is the US2 site;
wherein the first heterologous nucleic acid encodes an Infectious Bursal Disease Virus viral protein 2 (IBDV VP2), an Infectious Laryngotracheitis Virus glycoprotein D (ILTV gD) and an Infectious Laryngotracheitis Virus glycoprotein I (ILTV gI), and wherein the second heterologous encodes a Newcastle Disease Virus fusion protein (NDV F); and
wherein the first heterologous nucleic acid is configured to be transcribed in the opposite direction relative to a UL54.5 gene of the rHVT.
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