	US 11,730,135 B2
	*Resistance in plants of Solanum lycopersicum* to the tobamovirus tomato brown rugose fruit virus**
Varda Ashkenazi, Berurim (IL); Yaniv Rotem, Berurim (IL); Ron Ecker, Berurim (IL); Shai Nashilevitz, Berurim (IL); and Naama Barom, Berurim (IL)
Assigned to VILMORIN & CIE., Paris (FR)
Filed by VILMORIN & CIE, Paris (FR)
Filed on Nov. 25, 2019, as Appl. No. 16/694,089.
Application 16/694,089 is a continuation of application No. PCT/IB2019/000674, filed on Jun. 14, 2019.
Application PCT/IB2019/000674 is a continuation in part of application No. PCT/EP2018/064055, filed on May 29, 2018.
Claims priority of application No. 17305644 (EP), filed on Jun. 1, 2017.
Prior Publication US 2020/0077614 A1, Mar. 12, 2020
This patent is subject to a terminal disclaimer.
Int. Cl. A01H 5/10 (2018.01); A01H 6/82 (2018.01); C12Q 1/6895 (2018.01); C12Q 1/6827 (2018.01)

CPC A01H 6/825 (2018.05) [A01H 5/10 (2013.01); C12Q 1/6827 (2013.01); C12Q 1/6895 (2013.01); C12Q 2600/13 (2013.01)]

20 Claims

1. A Solanum lycopersicum plant resistant to Tomato Brown Rugose Fruit virus (TBRFV) comprising in its genome the combination of:

a) the Tm-1 resistance gene on chromosome 2, and

b) at least one quantitative trait locus (QTL) chosen from QTL3 on chromosome 11, QTL1 on chromosome 6 and QTL2 on chromosome 9, that independently confer to the plant foliar and/or fruit tolerance to TBRFV, wherein said QTLs are present in the genome of a plant, seeds of which designated HAZTBRFVRES1 have been deposited with NCIMB under accession number 42758,

wherein said QTLs are to be found, for QTL1, on chromosome 6, within the chromosomal region delimited by TO-0005197 (SEQ ID NO:1) and TO-015581 (SEQ ID NO:2), for QTL2, on chromosome 9, within the chromosomal region delimited by TO-0180955 (SEQ ID NO:3) and TO-0196109 (SEQ ID NO:6) and for QTL3, on chromosome 11, within the chromosomal region delimited by TO-0122252 (SEQ ID NO:7) and TO-0162427 (SEQ ID NO:18).